Druid
Magic
Druid
Upgrades
Path 1
1. Hard Thorns$350total $350Hard thorns can pop 2 Bloons each and pop any Bloon type.
2. Heart of Thunder$850total $1 200Unleashes regular blasts of forked lightning that arc from Bloon to Bloon.
3. Druid of the Storm$1 700total $2 900Gusts of wind blow Bloons off the track away from the exit.
4. Ball Lightning$4 500total $7 400Able to perceive Camo Bloons. Creates powerful balls of lightning that can freeze Bloons. Additional lightning bolts arc off main attack to nearby Bloons.
5. Monarch of Storms$60 000total $67 400Calls forth a relentless barrage of superstorms. Blasts all Bloon types for massive damage and blows them away from the exit.
Path 2
1. Thorn Swarm$250total $250Shoots 8 thorns per shot instead of 5.
2. Heart of Oak$350total $600Attacks convert Regrow Bloons into normal Bloons. Gains 1% pierce for every life gained from the starting value after the upgrade. Maxes at 100% and diminishes if lives are lost.
3. Druid of the Jungle$1 050total $1 650Calls a vine from the ground to entangle and crush Bloons one at a time, leaving behind a deadly thorn pile.
4. Jungle's Bounty$4 900total $6 550Vines create stronger thorn piles. Druid generates cash and lives at the end of each round, plus extra cash per Banana Farm near the Druid. Vine Crush Ability: Vines grab many Bloons at once, following Druid's target priority.
5. Spirit of the Forest$35 000total $41 550Grows thorned vines along the path that deal constant damage and bonus damage to ceramics. Vines nearest the Spirit of the Forest do more damage. Activated ability grabs more Bloons.
Path 3
1. Druidic Reach$100total $100Increases range by a large amount.
2. Heart of Vengeance$300total $400Gains 10% attack speed and 1% for every life lost from the starting value after the upgrade. Maxes at 50% and diminishes if lives are restored.
3. Druid of Wrath$600total $1 000Gains attack speed during the round as long as the Druid is attacking Bloons.
4. Poplust$2 350total $3 350Boosts attack speed and pierce for other Druids in the radius. Can stack up to 5 times on any single Druid.
5. Avatar of Wrath$45 000total $48 350The more Bloons there are, the more damage it does!
Paragon
Root of all Nature$475 000That from which all life branches.
Abilities
| Ability | From | Cooldown | Effect | Description |
|---|---|---|---|---|
 Jungle's Bounty | 0-4-0: Jungle's Bounty | 0 s | – | |
 Vine Crush | 0-4-0: Jungle's Bounty | 30 s | radius 4; damage 9 999 999 | Vine_Crush_Description |
| Primordial Wrath | Paragon: Root of all Nature | 9 s | – | Primordial Wrath Ability |
Crosspath
Pick the path tiers like in the game. Shown: the cost with all upgrades, attacks, cooldown, damage and pierce of every projectile, nested ones included (explosions, fragments).
Numbers by tier
| Tier | Range | Cooldown, s | Damage per shot | Pierce | Rough DPS | Cannot pop |
|---|---|---|---|---|---|---|
| 000 | 35 | 1.1 | 5 | 1 | 4.5455 | Lead Frozen |
| 100 | 35 | 1.1 | 5 | 2 | 4.5455 | – |
| 200 | 35 | 1.1 | 5 | 2 | 4.5455 | – |
| 300 | 35 | 1.1 | 5 | 2 | 4.5455 | – |
| 400 | 35 | 1.1 | 5 | 2 | 4.5455 | – |
| 500 | 35 | 4 | 100 | 200 | 25 | – |
| 010 | 35 | 1.1 | 8 | 1 | 7.2727 | Lead Frozen |
| 020 | 35 | 1.1 | 8 | 1 | 7.2727 | Lead Frozen |
| 030 | 35 | 2.6 | 9 999 999 | 9 999 999 | 3846153.4615 | Lead Frozen |
| 040 | 45 | 2.6 | 9 999 999 | 9 999 999 | 3846153.4615 | Lead Frozen |
| 050 | 45 | 2.6 | 9 999 999 | 9 999 999 | 3846153.4615 | Lead Frozen |
| 001 | 45 | 1.1 | 5 | 1 | 4.5455 | Lead Frozen |
| 002 | 45 | 1.1 | 5 | 1 | 4.5455 | Lead Frozen |
| 003 | 45 | 1.1 | 5 | 1 | 4.5455 | Lead Frozen |
| 004 | 45 | 1.1 | 5 | 1 | 4.5455 | Lead Frozen |
| 005 | 50 | 0.55 | 20 | 1 | 36.3636 | Lead Frozen |
| Paragon | 50 | 0.5 | 490 | 3 | 980 | – |
Knowledge
| Knowledge | Effect | Works from |
|---|---|---|
| Hot Magic | All Magic type towers can pop Frozen Bloons. | placement |
| Lingering Magic | Longer projectile lifespan for Wizard Monkey, Super Monkey, Skywarden, and Druid. | placement |
| Tiny Tornadoes | Druid tornadoes spawn 3 smaller tornadoes when they expire. | placement |
| Warm Oak | Heart of Oak cost reduced by 100. | placement |
| Vine Rupture | Grants Spirit of the Forest Druid a new ability to burst all track vines, doing big damage to all non-Lead Bloons. New vines will regrow after each Rupture. | 0-5-0 |
| Cold Front | Ball Lightning's additional lightning bolts also freeze Bloons. | 4-0-0 |
Plus the ones every tower gets:
From game data: the tower model names the knowledge, or the knowledge names the tower or one of its upgrades. "Works from" lists the first tiers whose model has it.